Paul Tremelling joined the Royal Navy in 1996 and flew the Sea Harrier on 800, 801 and 899 Naval Air Squadrons. Qualifying as an Air Warfare Instructor, he transitioned to the Harrier GR9 and served on the Naval Strike Wing, a unit where he became Senior Pilot. Selected for exchange duties with the USN, Tremelling flew F/A-18E Super Hornet with VFA-25, The Fist of The Fleet. He fitted in three desk tours, two Battle Staff jobs and operational duties in Afghanistan and off Libya. He was Mentioned in Despatches following an action fought in Afghanistan.